Distribution of Potamophylax cingulatus. Distribution in 1974–1978 (shaded). More recent records (+) and survey sites where the species was not recorded (o). Glaciers shown in grey. Shortest distance shown from the 1974–1978 distribution to the sites of first record in the 2005–2006 studies. Dispersal rate are 190 km/28 years = 7 km/year, 150 km/16 years = 9 km/year and 175 km/19 years = 9 km/year.